The PCIe M.2 SSD would only be able to operate at PCIe x2 (2-lane functionality) speeds within that motherboard. If you purchase a motherboard that supports PCIe x4 speeds, your x4-capable M.2 SSD should work as expected within that environment. In addition, there are PCIe limitations on system boards where the total number of PCIe lanes could be exceeded, limiting the PCIe M.2 x4 SSD to either have 2 lanes or even none.